Jamieson, B.G.M., Guinot, D. and Richer de Forges, B. 1993l. Spermatozoal ultrastructure in four genera of Homolidae (Crustacea, Decapoda): exemplified by Homologenus sp., Latreillopsis sp., Homolomannia sibogae and Paromolopsis boasi. Helgolander Meeresuntersuchungen 47, 323-334.

  The spermatozoa of Homologenus sp., Latreillopsis sp., Homolomannia sibogae
  and Paromolopsis boasi confirm characteristics of a distinctive homolid
  spermatozoon previously established for Homola sp., Paromola sp. and
  Paromola petterdi. Homolid features are (1) moderate anteroposterior
  depression of the acrosome (ratio of length: width 0.4-0.6) as in
  lyreidine raninids (0.5), depression being greater in dromiids and
  dynomenids (both 0.3); (2) the capitate form of the perforatorium, shared
  with dromiids, dynomenids and lyreidine raninids; (3) the autapomorphic
  spiked-wheel form of the anterior expansion of the perforatorium; (4)
  horizontal zonation of the acrosome is possibly a unique synapomorphy of
  homolids with dromiids and dynomenids, and therefore an autapomorphy of
  the dromioid-homolid assemblage. In dromiids the posterior zone is
  proportionately the larger, while in homolids the anterior zone is the
  larger. The anterior zone is complexly subdivided in dynomenids; (5) the
  autapomorphic presence of numerous radial arranged extensions of the
  acrosomal operculum into the perforatorium; (6) presence of nuclear arms,
  a symplesiomorphy of all investigated crabs, but small or questionably
  sometimes absent in Dromiidae; (7) absence of microtubules from the
  nuclear arms, as in dromiids, raninids, higher heterotremes and
  thoracotremes; (8) transient presence of a posterior median process of the
  nucleus. The process is not seen in dromiids but occurs in anomurans and
  lower heterotremes; (9) apical perforation of the operculum, also seen,
  apparently symplesiomorphically, in dromiids, raninids, and lower
  heterotreme families; (10) absence of an acrosome ray zone, probably
  homoplasic with absence in raninids; (11) location of most of the
  cytoplasm, including tortuous membranes and degenerating mitochondria,
  below the acrosome, also seen in Lyreidus; (12) presence, in at least some
  species, of centrioles, unknown in dromiids and raninids and variable in
  occurrence in heterotremes.