Jamieson, B.G.M. 1994j. Phylogeny of the Brachyura with particular reference to the Podotremata: evidence from a review of spermatozoal ultrastructure (Crustacea, Decapoda). Philosophical Transactions of the Royal Society London, B 345, 373-393.
Parsimony analysis, whether using only spermatozoal characters orspermatozoal and non-spermatozoal characters, suggests that: (i) the   Brachyura is a monophyletic taxon relative to the outgroup, three   paguroids; (ii) the Podotremata is a monophyletic taxon and the sister   -group of the heterotreme-thoracotreme assemblage; (iii) within the   Podotremata, the Dromiidae appears paraphyletic as its clade includes   Paradynomene; (iv) the Homolidae, is a monophyletic family; (v) the   Raninoidea and Cyclodorippoidea are sister groups within a monophyletic   (unnamed) clade; (vi) Latreillia sp. forms a polytomy with Homolidae +   Raninoidea-Cyclodorippoidea; and (vii) within the heterotremethoracotreme   assemblage, the Thoracotremata is a monophyletic taxon but the   Heterotremata s. strict. is a paraphyletic grouping. Analyses based on   sperm data alone differ from the combined data in not including the   Homolidae in the Archaeobrachyura (Homolidae + Raninoidea   -Cyclodorippoidea), thus the Dromiidae-Paradynomene + Homolidae +   Latreillia form a polytomous clade which is the sister-group of the   Raninoidea + Cyclodorippoidea. Spermatozoal data also give majids the most   basal position in the Heterotremata whereas for the combined data   Neodorippe (a shell carrier) appears the least modified member of the   heterotreme-thoracotreme assemblage. These findings are viewed in the   perspective of other, notably molecular, studies. Spermatozoal   synapomorphies are listed and illustrated.