148. Jamieson, B.G.M., Oliver, S.C. and Scheltinga, D.M. 1996a. The ultrastructure of the spermatozoa of Squamata--I. Scincidae, Gekkonidae and Pygopodidae (Reptilia). Acta Zoologica 77, 85-100.
Squamate autapomorphies seen in sperm of the Scincidae (e.g. Ctenotus
  robustus, Carlia pectoralis, Cryptoblepharus virgatus, and Lampropholis
  delicata) are penetration of the fibrous sheath of the axoneme into the
  midpiece, and the paracrystalline subacrosomal cone. Sphenomorphus group
  spermatozoa (e.g. Ctenotus) and the Egernia group (Tiliqua) differ from
  the more derived Eugongylus group (C. virgatus, L. delicata and C.
  pectoralis) in that the acrosome is elongate and apically depressed; the
  perforatorium is strongly oblique; the midpiece is relatively short, with
  four dense ring structures in longitudinal succession; mitochondria are
  columnar; and enlarged peripheral fibres 3 and 8 do not show the gross
  anterior enlargement seen in Carlia and Lampropholis. Heteronotia binoei
  (Gekkonidae) sperm have no epinuclear electron-lucent region; nuclear
  shoulders are smooth, as in sphenomorph but not Eugongylus group skinks;
  mitochondria are columnar; unlike skinks, the median surfaces of the
  mitochondria are indented by triangular, sometimes longitudinally,
  interconnected dense bodies. In Lialis burtonis (Pygopodidae) sperm, the
  perforatorium extends virtually to the tip of the fore-shortened apically
  domed acrosome; nuclear shoulders are absent; the mitochondria alternate
  singly or in groups with one or more dense bodies which also form an
  interrupted collar around the distal centriole. Spermatozoal
  ultrastructure suggests that a common ancestry of snakes and pygopods
  deserves consideration.